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<article xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:ali="http://www.niso.org/schemas/ali/1.0/" dtd-version="1.3" article-type="research-article" xml:lang="en"><front><journal-meta><journal-id journal-id-type="publisher-id">fg</journal-id><journal-title-group><journal-title>Forum Geografi</journal-title><abbrev-journal-title abbrev-type="publisher">fg</abbrev-journal-title></journal-title-group><issn pub-type="ppub">0852-0682</issn><issn pub-type="epub">2460-3945</issn><publisher><publisher-name>Universitas Muhammadiyah Surakarta</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="publisher-id">15980</article-id><title-group><article-title>Linking Weather Variability and Wildfires Across Tropical Ecoregions:  Evidence from Sumatra, Indonesia</article-title></title-group><contrib-group><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0009-0000-3462-5568</contrib-id><name><surname>Naim</surname><given-names>Asshaffa</given-names></name><xref ref-type="aff" rid="AFF-1"/></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0003-3497-0905</contrib-id><name><surname>Sekaranom</surname><given-names>Andung Bayu</given-names></name><xref ref-type="aff" rid="AFF-2"/><xref ref-type="corresp" rid="cor-0"/></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0002-6243-1346</contrib-id><name><surname>Sudrajat</surname><given-names>Sudrajat</given-names></name><xref ref-type="aff" rid="AFF-2"/></contrib><contrib contrib-type="author"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0001-8024-0845</contrib-id><name><surname>Nucifera</surname><given-names>Fitria</given-names></name><xref ref-type="aff" rid="AFF-3"/></contrib></contrib-group><aff id="AFF-1"><institution>Master in Geography Program, Faculty of Geography, Universitas Gadjah Mada, 55281, Yogyakarta</institution><country>Indonesia</country></aff><aff id="AFF-2"><institution>Department of Environmental Geography, Faculty of Geography, Universitas Gadjah Mada, 55281, Yogyakarta</institution><country>Indonesia</country></aff><aff id="AFF-3"><institution>Division of Sustainable Energy and Environment, Graduate School of Engineering, The University of Osaka, 565-0871, Osaka</institution><country>Japan</country></aff><author-notes><corresp id="cor-0">Corresponding author: Andung Bayu Sekaranom, Department of Environmental Geography, Faculty of Geography, Universitas Gadjah Mada, 55281, Yogyakarta, Indonesia. Email: <email>andung.geo@ugm.ac.id</email></corresp></author-notes><pub-date date-type="pub" publication-format="electronic" iso-8601-date="2026-7-6"><day>6</day><month>7</month><year>2026</year></pub-date><pub-date date-type="collection" publication-format="electronic" iso-8601-date="2026-7-6"><day>6</day><month>7</month><year>2026</year></pub-date><volume>40</volume><issue>3</issue><fpage>391</fpage><lpage>406</lpage><abstract><p>Wildfires in tropical regions exhibit strong spatial variability that reflects differences in ecoregion characteristics and weather sensitivity. This study aims to examine the relationship between weather dynamics and wildfire distribution across six different ecoregion characteristics in Sumatra. MODIS Fire hotspot data were used to analyze wildfire distribution from 2016–2023, combined with ERA5 and SMAP soil moisture data. Negative binomial regression and incidence rate ratio analysis were performed to obtain a statistical measure of the influence of weather. The results show that wildfire occurrence was highly concentrated in coastal and swamp ecoregions, with peat swamp forests (PSFs) accounting for the highest hotspot density. Pre-fire weather analyses showed that swamp and mangrove ecoregions exhibited higher sensitivity to relatively small rainfall deficits and temperature increases compared to upland forests. The negative binomial regression indicated that the strongest weather–wildfire associations were in freshwater swamp forest (FSFs), where weather variables yielded the highest pseudo-R² (0.226). The distinct weather sensitivity of each ecoregion was further explained by the IRR values, which showed strong temperature sensitivity in the Sunda Shelf mangrove (SSM) region and a prominent rainfall-related suppression of wildfire activity in FSFs. The findings demonstrate ecoregion-specific wildfire sensitivity and the need for targeted wildfire management strategies, particularly in peatland and coastal ecosystems in Sumatra.</p></abstract><kwd-group kwd-group-type="author-generated"><kwd>ecoregion</kwd><kwd>wildfire</kwd><kwd>Sumatra</kwd><kwd>precipitation</kwd><kwd>temperature</kwd><kwd>soil moisture</kwd></kwd-group><history><date date-type="received" iso-8601-date="2026-2-24"><day>24</day><month>2</month><year>2026</year></date><date date-type="rev-recd" iso-8601-date="2026-7-1"><day>1</day><month>7</month><year>2026</year></date><date date-type="accepted" iso-8601-date="2026-7-1"><day>1</day><month>7</month><year>2026</year></date></history><permissions><copyright-statement>Copyright © 2026 Asshaffa Naim, Andung Bayu Sekaranom, Sudrajat Sudrajat, Fitria Nucifera</copyright-statement><copyright-year>2026</copyright-year><copyright-holder>Asshaffa Naim, Andung Bayu Sekaranom, Sudrajat Sudrajat, Fitria Nucifera</copyright-holder><license xlink:href="https://creativecommons.org/licenses/by/4.0"><license-p>This article is distributed under the terms of the license at https://creativecommons.org/licenses/by/4.0.</license-p></license></permissions></article-meta></front><body><sec id="sec-1"><title>1. Introduction</title><p>Global warming has increased wildfire vulnerability by increasing temperatures and reducing precipitation levels, a situation which creates dry and flammable environmental conditions (<xref ref-type="bibr" rid="bib68">Gincheva et al., 2024</xref>). Wildfires, such as those in Australia during the 2019-2020 season, have burned approximately 3-4% of the world's land area, causing extensive ecosystem damage and producing large amounts of greenhouse gas emissions (<xref ref-type="bibr" rid="bib58">Chen &amp; Liu, 2024</xref>; <xref ref-type="bibr" rid="bib82">Nolan et al., 2020</xref>). These incidents are driven by a combination of dry weather conditions and human activities, such as land clearing, agriculture and deforestation (<xref ref-type="bibr" rid="bib77">Liu et al., 2024</xref>). Weather dynamics, particularly declining precipitation and rising temperatures, have been known to increase the number of wildfires by facilitating evapotranspiration and reducing fuel moisture (<xref ref-type="bibr" rid="bib59">Cho et al., 2024</xref>; <xref ref-type="bibr" rid="bib102">Wasserman &amp; Mueller, 2023</xref>). Several studies have shown that the influence of weather variability on wildfire activity is strongly mediated by the characteristics of soil, vegetation, hydrology and climate within an ecoregion (<xref ref-type="bibr" rid="bib81">Moradi &amp; Rahmati, 2023</xref>). Consequently, the distribution patterns vary across ecoregion types, making ecoregion-based analysis essential for understanding the mechanisms of wildfire occurrence and providing an ecologically grounded framework for understanding ecological sensitivity to wildfire. Addressing this variability requires spatially explicit evidence and analytical approaches that can capture related heterogeneity.</p><p>Satellite products such as MODIS and VIIRS enable near real-time detection of fire-prone locations through hotspot identification based on thermal anomaly algorithms (<xref ref-type="bibr" rid="bib57">Briones-Herrera et al., 2020</xref>). These hotspot datasets serve as proxies for wildfire occurrence, which, when combined with weather and soil conditions, allow for a comprehensive assessment of sensitivity across ecoregions. The use of negative binomial regression further strengthens analysis by quantifying the relationship between wildfire frequency and weather dynamics, while addressing overdispersion in hotspot data. By integrating satellite-based hotspot detection with reanalysis of climate and soil moisture data through an ecoregion-based statistical framework, this study not only contributes to advancing scientific understanding of wildfire–weather interactions in tropical landscapes, but also points to practical implications for early warning systems and management policies. Integrating ecoregion-specific weather sensitivity into prevention planning can enhance both short-term preparedness and long-term adaptation, thereby reducing ecological degradation and public health risks associated with recurrent wildfires.</p><p>Previous tropical wildfire studies have predominantly focused on large-scale climate variability, such as ENSO and IOD, as primary drivers of interannual wildfire variability across Southeast Asia (<xref ref-type="bibr" rid="bib56">Brasika, 2022</xref>; <xref ref-type="bibr" rid="bib83">Novitasari et al., 2019</xref>; <xref ref-type="bibr" rid="bib84">Nurdiati et al., 2022</xref>). Subsequent studies have incorporated local hydrological controls, peatland characteristics, and human-induced landscape modifications to improve risk assessment (<xref ref-type="bibr" rid="bib96">Salmayenti et al., 2025</xref>; <xref ref-type="bibr" rid="bib101">Taufik et al., 2022</xref>). Recently, machine learning approaches and fire danger models have been applied to evaluate spatial patterns of susceptibility under varying climatic conditions (<xref ref-type="bibr" rid="bib69">Hayasaka, 2023</xref>; <xref ref-type="bibr" rid="bib90">Prasetyo et al., 2022</xref>), while broader studies have documented the emergence of novel fire regimes across tropical forests worldwide (<xref ref-type="bibr" rid="bib76">Li et al., 2025</xref>; <xref ref-type="bibr" rid="bib87">Pacuk et al., 2025</xref>). Despite these advances, the role of ecoregional characteristics in shaping wildfire responses to short-term weather variability remains insufficiently understood. Previous studies generally focus on specific ecosystem, particularly peatlands, or assess fire-weather relationships at broad spatial scales without considering ecological heterogeneity among landscapes. Consequently, it remains unclear whether similar short-term weather anomalies exert comparable influences on wildfire occurrence across different ecoregions, or whether ecological characteristics modulate the magnitude and direction of these relationships (<xref ref-type="bibr" rid="bib92">Quan et al., 2023</xref>). This knowledge gap limits the ability to identify ecosystem-specific drivers and constrains the development of differentiated wildfire mitigation strategies. </p><p>To address this gap, this study was conducted to investigate wildfire-triggering weather conditions from an ecoregion perspective. Rather than treating tropical landscapes as environmentally homogeneous, the study examines whether the sensitivity of wildfire occurrence to short-term pre-fire weather variability differs systematically among distinct ecoregion types in Sumatra. By integrating MODIS FIRMS hotspot observations, ERA5 weather variables, and SMAP soil moisture data within an ecoregion-based statistical framework, the results quantify and compare fire-weather relationships across multiple tropical ecosystems simultaneously. Therefore, the study aims to 1) describe the spatial and temporal distribution of hotspots based on ecoregions; 2) analyze the spatial and temporal dynamics of precipitation, temperature and soil moisture before the emergence of hotspots; and 3) assess the relationship between hotspots and variability in precipitation, temperature and soil moisture. This approach will provide new insights into how ecological characteristics mediate wildfire responses to weather anomalies, advancing a more ecosystem-specific understanding of wildfire processes in tropical regions.</p></sec><sec id="sec-2"><title>2. Methods </title><p>The study framework comprised four sequential stages: data acquisition, data pre-processing, spatial and temporal analysis, and statistical analysis. A detailed workflow is presented in Figure <xref ref-type="fig" rid="fig-1">1</xref>. The methodological approach shared conceptual similarities with recent wildfire–weather analyses in other tropical and subtropical regions, yet differed in several key respects. For instance, studies in semi-arid tropical regions have used monthly or seasonal weather aggregates (<xref ref-type="bibr" rid="bib72">Justino et al., 2023</xref>), while our study used daily pre-fire windows to capture short-term atmospheric triggers. In addition, unlike analyses that treat entire islands as single analytical units (<xref ref-type="bibr" rid="bib84">Nurdiati et al., 2022</xref>), this study stratified observations by ecoregion to enable finer ecological differentiation. The integration of SMAP soil moisture alongside ERA5 data also distinguishes this framework from other studies which have relied solely on precipitation-based indices to address the need for multivariable fire risk modelling (<xref ref-type="bibr" rid="bib74">Krueger et al., 2022</xref>; <xref ref-type="bibr" rid="bib92">Quan et al., 2023</xref>). </p><fig id="fig-1"><label>Figure 1</label><caption><title>Study Framework (Source: Methodological Review).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83078"/></fig><sec id="sec-2_1"><title>2.1. Study Area</title><p>The study area was the island of Sumatra in Indonesia, with a total area of 448,975 km<sup>2</sup>, spanning 95°0′35′′ to 106°11′22′′ east and 5°56′36′′ south to 6°4′37′′ north. The island stretches from northwest to southeast, and is located southwest of the Malay Peninsula, bordered by the Indian Ocean to the west and east, and the Malacca Strait and South China Sea to the east. Topographically, it is characterized by mountainous terrain in its western and northern regions, while the eastern region is dominated by extensive lowlands. In addition, it has a tropical climate with two main seasonal patterns: an equatorial type with two rainfall peaks in the northern and central parts; and a monsoonal type with a single rainfall peak in the southern region (<xref ref-type="bibr" rid="bib53">Ariska et al., 2024</xref>). Figure <xref ref-type="fig" rid="fig-2">2</xref> shows that the study area covers six ecoregions based on biogeographical and ecological characteristics proposed by <xref ref-type="bibr" rid="bib62">Dinerstein et al. (2017)</xref>, consisting of tropical pine forest (TPF), montane rainforest (MRF), lowland rainforest (LRF), freshwater swamp forest (FSF), peat swamp forest (PSF), and Sunda Shelf mangrove (SSM) areas. </p><fig id="fig-2"><label>Figure 2</label><caption><title>Study Area (Source: Data Processing in QGIS 3.34).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83079"/></fig><p>Sumatra is one of the most critical regions in Indonesia for study of the link between weather variability and wildfire hotspots across ecoregions. Over the past decade, more than 2.8 million ha of land have burned on the island (<xref ref-type="bibr" rid="bib73">Kementerian Kehutanan, 2025</xref>). Sumatra hosts the largest peatland area in the country, covering approximately 6.4 million hectares, primarily in Riau, Jambi and South Sumatra (<xref ref-type="bibr" rid="bib94">Ritung et al., 2018</xref>). Peatland is highly vulnerable to wildfires because of its high organic content, which becomes extremely flammable under dry conditions, especially during prolonged droughts often triggered by global climate phenomena such as El Niño (<xref ref-type="bibr" rid="bib69">Hayasaka, 2023</xref>). Sumatra is also home to endangered and endemic species, which encounter heightened threats from recurring wildfires. Evidence shows that species diversity indices in burned forests are significantly lower than in unburned areas (<xref ref-type="bibr" rid="bib100">Syaufina et al., 2018</xref>). Due to this combination of weather sensitivity and ecological importance, Sumatra provides a critical setting for analyzing how weather variability shapes wildfire occurrence across ecoregions. </p><p>The study spanned eight years (2016-2023), a period selected to capture significant interannual climate variability, including both relatively wet and dry years, and to ensure temporal consistency across all datasets. The period was selected for several other reasons. First, the SMAP satellite became fully operational in April 2015, making 2016 the first complete calendar year with consistently available soil moisture data across all three datasets. Second, the period comprised marked interannual climate variability, including strong El Niño years and near-normal conditions, which allowed robust statistical analysis. Finally, the MODIS and ERA5 data for 2024 onwards were incompletely processed in the accessible repositories at the time of the analysis. </p></sec><sec id="sec-2_2"><title>2.2. Datasets</title><p>Four primary datasets were used, consisting of wildfire data, weather data, soil moisture data and ecoregion data. Each was downloaded using a common bounding box (7°S-7°N, 94°E-107°E) to ensure complete spatial coverage of the study area and consistent data processing. Wildfire data were in the form of fire hotspot locations, sourced from the MODIS Active Fire Product and accessed through NASA FIRMS (firms.modaps.eosdis.nasa.gov) (<xref ref-type="bibr" rid="bib67">Giglio et al., 2016</xref>). This dataset was generated from the identification of wildfire hotspots in MODIS imagery, based on a series of algorithms and thresholds that had been adjusted to local conditions (<xref ref-type="bibr" rid="bib67">Giglio et al., 2016</xref>). Each hotspot represented a thermal anomaly and indicated relative wildfire activity patterns rather than exact wildfire extent or perimeters. MODIS also provided a confidence value that represented the likelihood that a detected hotspot corresponded to an actual wildfire event. This feature served to filter the data and retain only high-confidence hotspots, which reduced false detections and improved the reliability of regional-scale wildfire pattern analysis.  </p><p>Weather data were represented by precipitation and temperature variables taken from the ERA5 datasets, a global weather parameter dataset that combines model output with observation data to provide temporally consistent climate variables, which is widely used to analyze long-term trends in climatic studies (<xref ref-type="bibr" rid="bib95">Sabljić et al., 2026</xref>). Daily total precipitation and 2m temperature data were obtained from the ERA5 reanalysis dataset (cds.climate.copernicus.eu) at a spatial resolution of 0.25° × 0.25° and hourly temporal resolution for the period 2016 to 2023 (<xref ref-type="bibr" rid="bib70">Hersbach et al., 2023</xref>). These variables were selected as fundamental drivers of wildfire risk. Temperature directly influences the rate of evapotranspiration, which dries out vegetation and other potential fuels, thereby leading to more flammable <xref ref-type="bibr" rid="bib102">conditions (Wasserman &amp; Mueller, 2023)</xref>. Precipitation is the primary source of moisture for both live and dead fuels, with its absence or scarcity being a direct indicator of drought conditions conducive to fire ignition and spread (<xref ref-type="bibr" rid="bib59">Cho et al., 2024</xref>; <xref ref-type="bibr" rid="bib75">Lafon &amp; Quiring, 2012</xref>).  </p><p>Soil moisture data were sourced from SMAP Surface and Root Zone Soil Moisture (appears.earthdatacloud.nasa.gov) at a spatial resolution of approximately 9 km and a 3-hourly temporal resolution (<xref ref-type="bibr" rid="bib93">Reichle et al., 2022</xref>). This variable was selected because it provides a more stable and integrated measure of landscape dryness than precipitation alone. In addition, it shows the cumulative effect of past rainfall and temperature, which indicates overall water stress on vegetation and the moisture content of deeper organic soil layers, crucial in peatland wildfire analysis (<xref ref-type="bibr" rid="bib74">Krueger et al., 2022</xref>). Building on this landscape-specific approach, the RESOLVE Ecoregions 2017 dataset was used to delineate distinct ecological zones for a stratified analysis (ecoregions.appspot.com) (<xref ref-type="bibr" rid="bib62">Dinerstein et al., 2017</xref>). The ecoregion represented broad biogeographical and ecological characteristics that were appropriate for capturing underlying climatic, hydrological and ecological controls, shaping long-term wildfire sensitivity across landscapes. While land use and land cover (LULC) change occurred, the ecoregion provided a more stable and ecologically meaningful basis than the dynamic land cover classes (<xref ref-type="bibr" rid="bib61">Curcan et al., 2023</xref>; <xref ref-type="bibr" rid="bib64">Drăguleasa et al., 2023</xref>; <xref ref-type="bibr" rid="bib79">Măceșeanu et al., 2026</xref>), especially given the island-wide scale of the study. </p></sec><sec id="sec-2_3"><title>2.3. Data Processing</title><p>All the variables were resampled to a common 0.25° grid corresponding to the ERA5 resolution using bilinear interpolation that estimated values at target grid points by computing a weighted average of the four nearest source grid cells to ensure spatial consistency across datasets. This resulted in approximately 2200 grid cells covering Sumatra. Temporally, ERA5 hourly data and SMAP 3-hourly data were aggregated to daily values by computing the daily means. Data processing and spatial analysis were conducted using QGIS version 3.34.12, while statistical analyses were performed using Python (including xarray, numpy and geopandas libraries) (<xref ref-type="bibr" rid="bib52">Arias-Muñoz et al., 2024</xref>; <xref ref-type="bibr" rid="bib90">Prasetyo et al., 2022</xref>). Raw hotspot data from MODIS were filtered to minimize false detections. Only hotspots with a confidence value &gt; 80% were retained, following the high-confidence threshold recommended in the product's user guide (<xref ref-type="bibr" rid="bib67">Giglio et al., 2016</xref>). Subsequently, these filtered points were aggregated to calculate fire density within the common grid. In addition to density calculations, the plotted hotspots were also classified based on their zones. Annual and monthly wildfire accumulation in each ecoregion was also calculated for temporal analysis. </p><p>For each hotspot, the corresponding weather and soil moisture data were extracted for the seven days preceding each hotspot detection, denoted as D-7 to D-0 to represent the pre-fire period. This window was selected based on previous studies that have consistently identified 5- to 10-day antecedent drying periods as the most critical precursors to fire ignition, with soil moisture and atmospheric dryness responding most strongly to rainfall deficits within this timeframe (<xref ref-type="bibr" rid="bib60">Chowdhury &amp; Hassan, 2015</xref>; <xref ref-type="bibr" rid="bib92">Quan et al., 2023</xref>). In addition, the lag time between rainfall cessation and critical near-surface moisture depletion in tropical peatlands have typically fallen within a 3-7 day range (<xref ref-type="bibr" rid="bib101">Taufik et al., 2022</xref>). This confirmed its suitability as the optimal temporal resolution for capturing short-term wildfire conditioning in the study context. The daily average value of each variable was calculated for each grid cell, while anomalies were calculated as the difference between the daily observed value and the pre-fire period mean across all hotspot events, as a standardized measure of departure from the pre-fire baseline condition. The processed hotspot densities and variables were analyzed on both monthly and annual scales. Data were classified and aggregated based on their ecoregion zones to facilitate a comparative analysis of wildfire dynamics. </p></sec><sec id="sec-2_4"><title>2.4. Statistical Analysis</title><p>Wildfire occurrences referred to count data that typically exhibited overdispersion, a scenario in which the variance in the data is significantly larger than the mean. This was common because many grid cells had zero wildfires, while a few exhibited a very high concentration. Therefore, to quantify the relationship between environmental conditions and wildfire occurrence, a negative binomial regression model was used. This statistical method was chosen to address wildfire data overdispersion. The appropriateness of the negative binomial model over Poisson regression was evaluated by testing the statistical significance of the dispersion parameter (α); a statistically significant α (p &lt; 0.05) indicated the presence of overdispersion.  The negative binomial model was a more robust and valid choice as it included an extra variable to account for this excess variance (<xref ref-type="bibr" rid="bib98">Stoklosa et al., 2022</xref>). The regression model was formally expressed as Equation 1:</p><p>where μᵢ represents the expected number of high-confidence hotspots in grid cell i; P, T and SM show the 7-day pre-fire mean values of precipitation (mm), temperature (°C) and soil moisture (%) respectively; β₀ is the intercept; β₁, β₂, and β₃ are the estimated regression coefficients; and εᵢ is the error term. The model incorporated a dispersion parameter α that allowed variance to exceed the mean, thereby addressing the overdispersion inherent in hotspot count data.  </p><p>The statistical significance of the predictor variables within the negative binomial regression was evaluated at a 95% confidence level (p-value &lt; 0.05). Variables yielding a p-value of less than 0.05 were considered statistically significant drivers of fire occurrence. In the model, the number of high-confidence hotspots per grid cell served as the dependent variable. The 7-day pre-fire averages of temperature, precipitation and soil moisture were used as the independent (predictor) variables, and the statistical analysis was performed for each ecoregion and for the whole region. Negative binomial analysis for the MRF and TPF ecoregions was conducted simultaneously because the number of hotspots in the TPF did not meet the minimum number of units. The results of this regression were presented in a table that included the pseudo-R2 variable, regression coefficients, and incidence rate ratio (IRR) values. Pseudo-R² measured the improvement in model fit of the full model relative to a null (intercept-only) model, in which higher values indicated a stronger association between the predictor variables and wildfire occurrence. Significance and coefficients indicated influential variables, while IRR described the magnitude of that influence on wildfire occurrence (<xref ref-type="bibr" rid="bib89">Piza, 2012</xref>). </p></sec></sec><sec id="sec-3"><title>3. Results</title><sec id="sec-3_1"><title>3.1. Spatial and Temporal Wildfire Distribution</title><p>The distribution of 19,450 hotspots recorded during the period 2016-2023 on Sumatra Island, as shown in Figure <xref ref-type="fig" rid="fig-3">3</xref>, revealed a concentration in the eastern region of the island, especially in the PSF ecoregion. Hotspots were aggregated within 0.25° grid cells, resulting in an average hotspot density of 28.85 points per grid cell. The maximum count within a single cell reached 1,222 hotspots, while 170 cells contained none. The highest number of hotspots was found in the PSF ecoregion, with 11,936 and a density of 13.7 per 100 km<sup>2</sup>. FSF and SSM exhibited high wildfire densities, with 9.7 and 6.6 wildfires per 100 km² respectively. Although the number of hotspots in LRF (4,679 hotspots) was the highest after PSF, the density was only 1.9 hotspots per 100 km<sup>2</sup>, which is lower than that of the freshwater swamp and mangrove ecoregions. The MRF and TPF ecoregions had the lowest density of hotspots, at 0.4 points per 100 km<sup>2</sup>.</p><fig id="fig-3"><label>Figure 3</label><caption><title>Sumatra Wildfire Hotspot Density Map (Source: MODIS Active Fire Data, Processed in QGIS 3.34).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83080"/></fig><p>The annual distribution of wildfires shown in Figure <xref ref-type="fig" rid="fig-4">4</xref> indicates significant spikes in dry years, particularly 2019 (11,241 hotspots) and 2023 (2,872 hotspots). Wildfire occurrence declined markedly after 2019, with relatively lower hotspot counts recorded during the period 2020 to 2022, before increasing again in 2023. Across all years, peat swamp forests (PSFs) consistently contributed the largest proportion of hotspots, particularly during high-fire years. In addition, Figure <xref ref-type="fig" rid="fig-5">5</xref> presents wildfire occurrences on the seasonal scale, which are strongly concentrated in the late dry season. Wildfire activities increased sharply in July and peaked between August and October throughout the study period. The highest monthly wildfire count was recorded in September (7,245 hotspots), followed by October (4,069 hotspots). However, wildfire occurrence remained comparatively low between December and June each year. A smaller secondary increase in hotspot activity was also observed from February to March. Similar to the annual distribution, PSF-dominated hotspot occurrence occurred during the peak wildfire months, while MRF and TPF consistently exhibited relatively low hotspot densities throughout the year. </p><fig id="fig-4"><label>Figure 4</label><caption><title>Annual Wildfire Hotspot Distribution Across Ecoregions in Sumatra (Source: MODIS Active Fire data).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83081"/></fig><fig id="fig-5"><label>Figure 5</label><caption><title>Monthly Wildfire Hotspot Distribution Across Ecoregions in Sumatra (Source: MODIS Active Fire Data).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83082"/></fig></sec><sec id="sec-3_2"><title>3.2. Pre-fire Weather Dynamics</title><p>Pre-fire weather dynamics in shown Figure <xref ref-type="fig" rid="fig-6">6</xref> demonstrate consistent patterns across ecoregions during the D−7 to D−0 period. The average decrease in rainfall over the 8 days was approximately 1.41 mm, with the decline becoming progressively more evident toward the wildfire occurrence date. Relative to the 8-day pre-fire average, most areas still exhibit positive rainfall anomalies from D-7 to D-5, particularly in the western part of Sumatra. A more pronounced decline was evident at D-4 and D-3, with mean daily rainfall anomalies across ecoregions ranging from −0.5 to 0 mm relative to the 8-day pre-fire baseline. This decreasing pattern continued across all ecoregions until D-1. Spatially, the onset of rainfall deficits occurred earlier in the eastern coastal plains, where negative anomalies were already evident at D-5, compared to the mountainous western regions. Referring to Table <xref ref-type="table" rid="table-1">1</xref>, the MRF ecoregion showed the greatest decrease in rainfall, with an average fall of 0.479 mm/day, while the FSF and PSF only experienced a decrease in rainfall of 0.071 mm/day and 0.183 mm/day. The absolute daily rainfall showed that MRF maintained the highest precipitation levels throughout the period, while FSF consistently recorded the lowest rainfall.  </p><fig id="fig-6"><label>Figure 6</label><caption><title>Pre-Fire Rainfall Dynamics Map (Source: MODIS Active Fire Data and ERA5 Total Precipitation Data, Processed in Python and QGIS 3.34).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83083"/></fig><table-wrap id="table-1"><label>Table 1</label><caption><title>Pre-fire Rainfall Dynamics.</title></caption><table frame="box" rules="all"><thead><tr><th rowspan="2"><p>Ecoregion</p></th><th colspan="8"><p>Rainfall (mm)</p></th></tr><tr><th><p>D-7</p></th><th><p>D-6</p></th><th><p>D-5</p></th><th><p>D-4</p></th><th><p>D-3</p></th><th><p>D-2</p></th><th><p>D-1</p></th><th><p>D-0</p></th></tr></thead><tbody><tr><td><p>FSF</p></td><td><p>1.28</p></td><td><p>1.21</p></td><td><p>1.02</p></td><td><p>0.88</p></td><td><p>1.07</p></td><td><p>1.09</p></td><td><p>0.70</p></td><td><p>0.78</p></td></tr><tr><td><p>LRF</p></td><td><p>3.66</p></td><td><p>3.28</p></td><td><p>2.98</p></td><td><p>2.62</p></td><td><p>2.10</p></td><td><p>1.75</p></td><td><p>1.34</p></td><td><p>1.59</p></td></tr><tr><td><p>MRF</p></td><td><p>5.60</p></td><td><p>5.36</p></td><td><p>3.98</p></td><td><p>3.74</p></td><td><p>3.38</p></td><td><p>2.61</p></td><td><p>2.48</p></td><td><p>2.25</p></td></tr><tr><td><p>PSF</p></td><td><p>2.39</p></td><td><p>2.11</p></td><td><p>1.62</p></td><td><p>1.48</p></td><td><p>1.30</p></td><td><p>1.38</p></td><td><p>0.99</p></td><td><p>1.11</p></td></tr><tr><td><p>TPF</p></td><td><p>2.65</p></td><td><p>2.55</p></td><td><p>0.91</p></td><td><p>1.00</p></td><td><p>1.51</p></td><td><p>0.82</p></td><td><p>0.51</p></td><td><p>0.90</p></td></tr><tr><td><p>SSM</p></td><td><p>3.72</p></td><td><p>3.95</p></td><td><p>3.17</p></td><td><p>2.93</p></td><td><p>2.30</p></td><td><p>2.09</p></td><td><p>1.55</p></td><td><p>2.27</p></td></tr></tbody></table></table-wrap><p><italic>FSF = freshwater swamp forest, LRF = lowland rainforest, MRF = montane rainforest, PSF = peat swamp forest, TPF = tropical pine forest, SSM = Sunda Shelf mangrove</italic></p><p>The pre-fire temperature dynamics shown in Figure <xref ref-type="fig" rid="fig-7">7</xref> indicate an increasing trend across all ecoregions, with absolute temperature values demonstrating stratification among ecoregions. FSF, MRF and TPF maintained lower temperatures, while LRF, PSF and SSM consistently exhibited higher ones. During the D-7 to D-5 period, most areas showed negative anomalies, especially in MRF and some parts of LRF, which had relatively stable pre-fire temperatures during the early observation period. The earliest and most widespread positive temperature anomaly appeared in the LRF and PSF ecoregions around D-4. Temperature increases became more evident from D-4 to D-2, with PSF recording a mean increase of 0.10°C over this 2-day interval. Meanwhile, the highest anomalies in D-1 to D-0 were more dominant in LRF. Referring to the values in Table <xref ref-type="table" rid="table-2">2</xref>, the rate of pre-fire temperature increase in PSF was comparatively lower (0.061°C/day) than in LRF (0.081°C/day), which represented a temperature rise of 0.57°C over the 7-day pre-fire window. The temperature change showed lower rates of increase in PSF (0.0610°C/day) and SSM (0.0639°C/day) than in MRF (0.0776°C/day) and TPF (0.1168°C/day). TPF, which had the highest temperature increase rate, showed an abrupt temperature increase of 0.62°C between D-2 and D-1, relative to the more gradual warming rates in other ecoregions.  </p><fig id="fig-7"><label>Figure 7</label><caption><title>Pre-fire Temperature Dynamics Map (Source: MODIS Active Fire Data and ERA5 2m-temperature Data, Processed in Python and QGIS 3.34).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83084"/></fig><table-wrap id="table-2"><label>Table 2</label><caption><title>Pre-fire Temperature Dynamics.</title></caption><table frame="box" rules="all"><thead><tr><th rowspan="2"><p>Ecoregion</p></th><th colspan="8"><p>Temperature (°C)</p></th></tr><tr><th><p>D-7</p></th><th><p>D-6</p></th><th><p>D-5</p></th><th><p>D-4</p></th><th><p>D-3</p></th><th><p>D-2</p></th><th><p>D-1</p></th><th><p>D-0</p></th></tr></thead><tbody><tr><td><p>FSF</p></td><td><p>28.32</p></td><td><p>28.30</p></td><td><p>28.41</p></td><td><p>28.43</p></td><td><p>28.49</p></td><td><p>28.44</p></td><td><p>28.53</p></td><td><p>28.52</p></td></tr><tr><td><p>LRF</p></td><td><p>27.09</p></td><td><p>27.17</p></td><td><p>27.21</p></td><td><p>27.34</p></td><td><p>27.37</p></td><td><p>27.48</p></td><td><p>27.55</p></td><td><p>27.66</p></td></tr><tr><td><p>MRF</p></td><td><p>21.72</p></td><td><p>21.74</p></td><td><p>21.80</p></td><td><p>21.88</p></td><td><p>21.96</p></td><td><p>22.09</p></td><td><p>22.14</p></td><td><p>22.27</p></td></tr><tr><td><p>PSF</p></td><td><p>28.15</p></td><td><p>28.17</p></td><td><p>28.28</p></td><td><p>28.35</p></td><td><p>28.44</p></td><td><p>28.45</p></td><td><p>28.53</p></td><td><p>28.59</p></td></tr><tr><td><p>TPF</p></td><td><p>20.23</p></td><td><p>20.43</p></td><td><p>20.35</p></td><td><p>20.36</p></td><td><p>20.43</p></td><td><p>20.37</p></td><td><p>20.99</p></td><td><p>21.04</p></td></tr><tr><td><p>SSM</p></td><td><p>27.82</p></td><td><p>27.81</p></td><td><p>27.85</p></td><td><p>27.96</p></td><td><p>28.01</p></td><td><p>28.09</p></td><td><p>28.17</p></td><td><p>28.25</p></td></tr></tbody></table></table-wrap><p><italic>FSF = freshwater swamp forest, LRF = lowland rainforest, MRF = montane rainforest, PSF = peat swamp forest, TPF = tropical pine forest, SSM = Sunda Shelf mangrove</italic><italic>.</italic></p><p>Figure <xref ref-type="fig" rid="fig-7">7</xref> shows a gradual change in pre-fire soil moisture anomalies  across most areas of Sumatra. Similar to the pattern shown by rainfall, the decrease in soil moisture during the period D-7 to D-5 was not very noticeable, although significant declines became evident closer to the wildfire occurrence date. Reductions in soil moisture intensified during the D-2 to D-0 window, with the highest absolute decrease in SSM (1.07%) and FSF (0.40%). Spatially, PSF, FSF and SSM exhibited higher soil moisture declines than other ecoregions, especially in the central part. However, LRF, MRF and TPF showed more modest declines, each decreasing by less than 0.40%. The LRF ecoregion showed a modest decrease, with an average pre-fire decline of approximately 1.05%. MRF and TPF also showed comparatively modest day-to-day soil moisture changes, ranging from 0.05 to 0.17% in one day. Table <xref ref-type="table" rid="table-3">3</xref> shows that FSF, PSF and SSM exhibited higher pre-fire soil moisture ranges of 54-55%, 65-67% and 75-77%, respectively, while the other three ecoregions exhibited similar lower soil moisture levels. Compared to the other ecoregions, swamp and mangrove ecosystems maintained higher baseline soil moisture despite experiencing noticeable short-term pre-fire declines.</p><fig id="fig-8"><label>Figure 8</label><caption><title>Pre-fire Soil Moisture Dynamics Map (Sources: MODIS Active Fire data and SMAP Soil Moisture Data Processed in Python and QGIS 3.34, 2025).</title></caption><graphic mimetype="image" mime-subtype="png" xlink:href="https://journals2.ums.ac.id/fg/article/download/15980/6342/83085"/></fig><table-wrap id="table-3"><label>Table 3</label><caption><title>Pre-fire Soil Moisture Dynamics.</title></caption><table frame="box" rules="all"><thead><tr><th rowspan="2"><p>Ecoregion</p></th><th colspan="8"><p>Soil Moisture (%)</p></th></tr><tr><th><p>D-7</p></th><th><p>D-6</p></th><th><p>D-5</p></th><th><p>D-4</p></th><th><p>D-3</p></th><th><p>D-2</p></th><th><p>D-1</p></th><th><p>D-0</p></th></tr></thead><tbody><tr><td><p>FSF</p></td><td><p>55.62</p></td><td><p>55.46</p></td><td><p>55.30</p></td><td><p>55.15</p></td><td><p>54.99</p></td><td><p>54.84</p></td><td><p>54.64</p></td><td><p>54.44</p></td></tr><tr><td><p>LRF</p></td><td><p>34.81</p></td><td><p>34.72</p></td><td><p>34.63</p></td><td><p>34.50</p></td><td><p>34.33</p></td><td><p>34.14</p></td><td><p>33.94</p></td><td><p>33.76</p></td></tr><tr><td><p>MRF</p></td><td><p>34.03</p></td><td><p>33.98</p></td><td><p>33.86</p></td><td><p>33.74</p></td><td><p>33.64</p></td><td><p>33.49</p></td><td><p>33.32</p></td><td><p>33.17</p></td></tr><tr><td><p>PSF</p></td><td><p>67.05</p></td><td><p>66.92</p></td><td><p>66.73</p></td><td><p>66.54</p></td><td><p>66.32</p></td><td><p>66.10</p></td><td><p>65.88</p></td><td><p>65.62</p></td></tr><tr><td><p>TPF</p></td><td><p>33.64</p></td><td><p>33.35</p></td><td><p>33.17</p></td><td><p>32.97</p></td><td><p>32.84</p></td><td><p>32.78</p></td><td><p>32.57</p></td><td><p>32.37</p></td></tr><tr><td><p>SSM</p></td><td><p>77.06</p></td><td><p>76.92</p></td><td><p>76.71</p></td><td><p>76.43</p></td><td><p>76.14</p></td><td><p>75.78</p></td><td><p>75.42</p></td><td><p>75.07</p></td></tr></tbody></table></table-wrap><p><italic>FSF = freshwater swamp forest, LRF = lowland rainforest, MRF = montane rainforest, PSF = peat swamp forest, TPF = tropical pine forest, SSM = Sunda Shelf mangrove</italic><italic>.</italic></p></sec><sec id="sec-3_3"><title>3.3. Wildfire and Weather Dynamics Linkage Based on Ecoregion</title><p>Negative binomial regression was applied to quantify the weather–wildfire relationship across ecoregions. The dispersion parameter (α) was greater than zero and statistically significant (α = 0.7184, p &lt; 0.05), which confirmed the presence of overdispersion and the appropriateness of this specification over Poisson regression. The relationship between the number of hotspots and rainfall, temperature and soil moisture, as presented in Table 4, shows varying results between ecoregions. The strongest relationship was shown by the FSF ecoregion, where the full model yielded the highest pseudo-R² (0.226), indicating the greatest improvement in model fit. Comparatively moderate fit improvements were observed in SSM (pseudo-R² = 0.140) and PSF (pseudo-R² = 0.052). Weaker fit improvement by weather variables was observed in LRF (pseudo-R² = 0.019) and in the combined MRF and TPF (pseudo-R² = 0.004). The coefficients of rainfall, temperature and soil moisture showed significant effects overall. However, there were variations in the effect with different levels of significance, both between ecoregions and between variables. In LRF and PSF, which had the largest number of wildfire hotspots, rainfall and temperature variables showed significant associations, while the soil moisture variable did not, demonstrating a significant statistical association in FSF and SSM instead. However, the effect was relatively modest, with a 1% increase in soil moisture contributing to a 1.97% increase in the expected incidence rate of wildfire hotspots in FSF and 3.41% in SSM. MRF and TPF exhibited no significant coefficients for any of the three weather variables.  </p><p>The IRR values show that a 1 mm increase in rainfall was associated with a 3.52% lower expected incidence rate of wildfire hotspots in LRF (IRR = 0.965) and a 10.35% lower expected incidence rate in PSF (IRR = 0.897). Meanwhile, a 1°C increase in temperature was associated with a 6.19% higher expected incidence rate of wildfire hotspots in LRF (IRR = 1.062) and an 11.45% higher rate in PSF (IRR = 1.115). The rainfall variable yielded a statistically significant coefficient in FSF, with the largest effect magnitude observed across all ecoregions. A 1 mm increase in rainfall was associated with a 13.52% lower expected incidence rate of wildfire hotspots in this ecoregion (IRR = 0.865). Temperature showed a strong association with wildfire occurrence in the SSM ecoregion, with an IRR value of 1.971. This indicates that a 1°C increase in temperature was associated with a 97.1% higher expected incidence rate of wildfire hotspots. However, this magnitude must be interpreted with caution, as the observed pre-fire temperature range in SSM spanned only approximately 0.43°C across the 7-day pre-fire window.  </p><table-wrap id="table-4"><label>Table 4</label><caption><title>Negative Binomial Regression Result.</title></caption><table frame="box" rules="all"><thead><tr><th rowspan="2"><p>Ecoregion</p></th><th rowspan="2"><p>No. Obs</p></th><th rowspan="2"><p>psd. R<sup>2</sup></p></th><th colspan="3"><p>Coefficient</p></th><th colspan="3"><p>IRR</p></th></tr><tr><th><p>rf</p></th><th><p>tm</p></th><th><p>sm</p></th><th><p>rf</p></th><th><p>tm</p></th><th><p>sm</p></th></tr></thead><tbody><tr><td><p>FSF</p></td><td><p>402</p></td><td><p>0.226</p></td><td><p>-0.145*</p></td><td><p>0.056</p></td><td><p>0.020*</p></td><td><p>0.865*</p></td><td><p>1.058</p></td><td><p>1.020*</p></td></tr><tr><td><p>LRF</p></td><td><p>2670</p></td><td><p>0.019</p></td><td><p>-0.036*</p></td><td><p>0.060*</p></td><td><p>0.002</p></td><td><p>0.965*</p></td><td><p>1.062*</p></td><td><p>1.002</p></td></tr><tr><td><p>MRF &amp; TPF</p></td><td><p>229</p></td><td><p>0.004</p></td><td><p>0.006</p></td><td><p>-0.006</p></td><td><p>0.013</p></td><td><p>1.006</p></td><td><p>0.994</p></td><td><p>1.013</p></td></tr><tr><td><p>PSF</p></td><td><p>2344</p></td><td><p>0.052</p></td><td><p>-0.109*</p></td><td><p>0.108*</p></td><td><p>0.001</p></td><td><p>0.897*</p></td><td><p>1.115*</p></td><td><p>1.001</p></td></tr><tr><td><p>SSM</p></td><td><p>216</p></td><td><p>0.140</p></td><td><p>-0.036</p></td><td><p>0.679*</p></td><td><p>0.034*</p></td><td><p>0.965</p></td><td><p>1.971*</p></td><td><p>1.034*</p></td></tr><tr><td><p>All</p></td><td><p>5548</p></td><td><p>0.166</p></td><td><p>-0.089*</p></td><td><p>0.109*</p></td><td><p>0.013*</p></td><td><p>0.915*</p></td><td><p>1.115*</p></td><td><p>1.014*</p></td></tr></tbody></table></table-wrap><p><italic>FSF = freshwater swamp forest, LRF = lowland rainforest, MRF = montane rainforest, PSF = peat swamp forest, TPF = tropical pine forest, SSM = Sunda Shelf mangrove</italic><italic>, </italic><italic>No. Obs = </italic><italic>number</italic><italic> of </italic><italic>observations</italic><italic>, </italic><italic>p</italic><italic>sd</italic><italic>.</italic><italic> R</italic><italic><sup>2</sup></italic><italic> = pseudo-R</italic><italic><sup>2</sup></italic><italic>, IRR = Incidence Rate Ratio, </italic><italic>rf</italic><italic> = </italic><italic>rainfall</italic><italic>, </italic><italic>tm</italic><italic> = </italic><italic>temperature</italic><italic>, </italic><italic>sm</italic><italic> = </italic><italic>soil</italic><italic>moisture</italic></p><p><italic>*Statistical significance at p &lt; 0.0</italic><italic>5</italic><italic>.</italic></p></sec></sec><sec id="sec-4"><title>4. Discussion</title><p>Our spatiotemporal analysis of wildfire dynamics across Sumatra reveals that the relationship between short-term weather variability and wildfire occurrence is strongly mediated by ecoregional characteristics, particularly hydrological conditions, ecosystem structure, and landscape disturbance. The spatial difference of wildfire concentration between FSF, PSF and SSM with LRF, MRF and TPF was consistent with recent global results showing that swamp and mangrove ecosystems are increasingly prone to wildfires due to hydrological disturbance, including altered water balance and vegetation stress associated with coastal drying during prolonged dry seasons (<xref ref-type="bibr" rid="bib58">Chen &amp; Liu, 2024</xref>; <xref ref-type="bibr" rid="bib85">O et al., 2020</xref>). Wildfire vulnerability is further intensified by the land-use conversion, peat drainage and deforestation associated with resource exploitation (<xref ref-type="bibr" rid="bib65">Eddy et al., 2021</xref>). These anthropogenic disturbances also increase the sensitivity of tropical ecosystems to short-term weather anomalies by altering hydrological conditions, lowering water tables, and increasing fuel flammability during dry periods (<xref ref-type="bibr" rid="bib96">Salmayenti et al., 2025</xref>; <xref ref-type="bibr" rid="bib101">Taufik et al., 2022</xref>). Although anthropogenic variables were not included in this study, the observed weather–wildfire relationships must be interpreted within the broader context of human-modified landscapes. For instance, the low density of hotspots in MRF and TPF suggests that these ecosystems are relatively more resistant to wildfire occurrence and less exposed to anthropogenic ignition pressures. This pattern can be seen in the mountainous topography of western Sumatra, with limited human disturbance and wildfire exposure (<xref ref-type="bibr" rid="bib54">Avcıoğlu et al., 2024</xref>). However, the reduced wildfire frequency in these highland ecosystems does not necessarily imply immunity. A study of tropical montane zones by <xref ref-type="bibr" rid="bib99">Swann et al. (2023)</xref> revealed that rising temperatures and precipitation shifts were extending wildfire seasons and altering wildfire boundaries, even in these stable ecosystems. </p><p>Temporally, the extreme wildfire years were consistent with large-scale ENSO–IOD influences referred to in the introduction. During El Niño events, suppressed rainfall prolongs dry season conditions into September–November, delaying fuel rewetting and extending the wildfire-prone period. The seasonal pattern of wildfired indicated that wildfire activity intensifies after the climatological peak of the dry season (June–August) in both equatorial and monsoonal climates. This seasonal pattern also shows the differing climatic regimes across Sumatra, in which the lower hotspot peak during February–March corresponds to the first dry season in equatorial climate regions. Meanwhile, the delayed responses of wildfire activity are consistent with a recent tropical study showing an inverse relationship between precipitation and wildfire occurrence, suggesting a time-lagged response of wildfire activity to rainfall and fuel drying conditions (<xref ref-type="bibr" rid="bib76">Li et al., 2025</xref>).  </p><p>The observed pre-fire weather dynamics consistently revealed a difference between coastal lowland and upland ecosystems. Coastal swamp ecoregions remain vulnerable to wildfire even under modest rainfall deficits, likely due to the rapid increase in fuel flammability associated with organic-rich surface layers and shallow water-table responses (<xref ref-type="bibr" rid="bib71">Irfan et al., 2021</xref>). On the other hand, mountainous areas experience higher rainfall that buffer them against short-term moisture stress. This pattern is similar to that of other humid tropical regions, where orographic precipitation and lower vapor pressure deficits confer greater climatic resilience (<xref ref-type="bibr" rid="bib75">Lafon &amp; Quiring, 2012</xref>; <xref ref-type="bibr" rid="bib88">Petrov et al., 2023</xref>). Temperature dynamics show a similar ecoregional contrast, including the progressive pre-fire warming observed across all ecoregions, consistent with global evidence that elevated temperatures increase the probability of ignition (<xref ref-type="bibr" rid="bib78">Ma et al., 2024</xref>). Meanwhile, the persistent thermal stratification, with FSF, PSF and SSM maintaining higher temperatures than MRF and TPF, reflect their contrasting latitudinal and elevational positions. In dense, moisture-rich tropical ecosystems, temperature effects are often manifested indirectly through enhanced evapotranspiration, atmospheric drying and reductions in canopy fuel moisture (<xref ref-type="bibr" rid="bib86">O’Donnell et al., 2011</xref>). Together, these atmospheric differences show that hydrologically-sensitive coastal ecosystems respond rapidly to short-term atmospheric drying than upland forest systems (<xref ref-type="bibr" rid="bib66">Flannigan et al., 2015</xref>).  </p><p>While rainfall and temperature primarily represent short-term atmospheric variability, soil moisture provides a more direct representation of hydrological conditions related to fuel availability and combustion persistence. The 7-day average soil moisture could not fully capture threshold-based or lagged hydrological effects underlying the abrupt ignition processes associated with critical moisture thresholds. However, such soil moisture deficits still reflect ignition potential and influence wildfire severity through their effect on fuel continuity and combustion completeness (<xref ref-type="bibr" rid="bib91">Pulla et al., 2016</xref>). In swamp-dominated regions with a high soil moisture baseline, a 5% decline corresponds to an exponential increase in smouldering depth and the overall hotspot number (<xref ref-type="bibr" rid="bib56">Brasika, 2022</xref>). Notably, ecoregions that exhibited larger pre-fire soil moisture declines, such as FSF, PSF and SSM, were also those with the highest hotspot densities. In these ecosystems, wildfire susceptibility is more appropriately understood as a function of relative drying from normally saturated baseline conditions, rather than of absolute moisture levels (<xref ref-type="bibr" rid="bib55">Azmi et al., 2025</xref>; <xref ref-type="bibr" rid="bib101">Taufik et al., 2022</xref>). This mechanism explains the counterintuitive positive coefficients of soil moisture in FSF and SSM. Although the negative binomial regression identified statistically significant associations, the corresponding IRR indicated relatively small changes (less than 5%), suggesting that soil moisture influences wildfire in a nonlinear and potentially threshold-based manner. The regression analysis quantitatively supported the descriptive spatiotemporal patterns observed across the six ecoregions, across which the Pseudo-R² values were relatively modest, indicating that the regression models captured only limited improvements in fit relative to null models within this complex wildfire system. The values suggest that meteorological variability alone accounted for only a proportion of the observed variation in wildfire occurrence, consistent with previous findings on tropical ecosystems, indicating that weather variables function more as short-term  enabling conditions than as sole determinants of wildfire activity (<xref ref-type="bibr" rid="bib87">Pacuk et al., 2025</xref>).  </p><p>The ecoregion-specific regression coefficients and IRR values quantitatively demonstrated significant differences in wildfire sensitivity between ecosystems. The results were consistent with previous studies showing that the relationship between rainfall and wildfire activity in humid tropical regions is highly nonlinear, where even short rainfall events rapidly suppress wildfire spread in swamp ecosystems (<xref ref-type="bibr" rid="bib80">Mishra &amp; Singh, 2010</xref>). Similarly, <xref ref-type="bibr" rid="bib97">Schaefer and&amp; Magi (2019)</xref>demonstrated that the sensitivity of wildfire occurrence to rainfall was particularly pronounced in areas with shallow water tables and organic-rich soils, such as FSF and PSF. The relatively large IRR associated with temperature in SSM was consistent with recent observations in coastal tropical environments, in which canopy structure and salt exposure were shown to intensify surface drying under warmer conditions (<xref ref-type="bibr" rid="bib63">Dookie et al., 2025</xref>). </p><p>The absence of significant coefficients in MRF and TPF was consistent with results from <xref ref-type="bibr" rid="bib88">Petrov et al. (2023)</xref>, indicating that montane and coniferous ecosystems typically exhibited higher climatic buffering due to lower human ignition pressure and more stable moisture regimes. Therefore, the weak weather sensitivity in these high-elevation systems demonstrates both ecological resistance and limited anthropogenic exposure. The difference between highly sensitive coastal ecosystems and comparatively stable upland systems emphasizes the importance of ecoregion-specific climatic thresholds in shaping wildfire susceptibility across Sumatra. In addition, rainfall and temperature were associated with short-term atmospheric conditions linked to wildfire occurrence, while soil moisture showed broader hydrological constraints on sustained combustion. The results emphasize that wildfire vulnerability in tropical landscapes is strongly conditioned by the interaction between short-term weather variability, hydrological sensitivity, and ecosystem-specific landscape characteristics. </p></sec><sec id="sec-5"><title>5. Conclusion</title><p>In conclusion, the study demonstrates that wildfire vulnerability in tropical landscapes is strongly differentiated by ecosystem type, with coastal wetland systems exhibiting the greatest sensitivity to short-term hydrometeorological variability. Reduced rainfall and elevated temperatures are consistently associated with increased wildfire occurrence, particularly in carbon-rich peatland and mangrove ecosystems. These results reinforce growing evidence from recent tropical wildfire studies that degraded tropical wetlands are highly sensitive to short-term hydrometeorological fluctuations. As climate change intensifies, the risk of recurrent wildfire in these ecosystems escalates, not only in Sumatra, but also across humid tropical regions. Recognizing ecoregion-specific responses to weather variability is essential for improving fire danger forecasting, early warning systems, and regionally targeted wildfire mitigation policies.   </p><p>Unlike many previous tropical wildfire studies that have focused primarily on peatlands or seasonal drought conditions, the integration of weather sensitivity into ecoregion-based wildfire assessment in this study demonstrates that weather–wildfire relationships vary significantly across ecoregions and are strongly mediated by ecosystem-specific hydrological characteristics. This can improve the spatial prioritization of wildfire prevention efforts applied to tropical landscapes and coastal ecosystems. From a practical perspective, the findings offer valuable insights to support environmental agencies and policymakers in identifying ecosystems that require more intensive hydrological monitoring and wildfire prevention during drought periods. However, the study is limited by its reliance on moderate-resolution satellite data and the use of relatively simple meteorological variables. Future studies should integrate finer-scale climate, hydrological, land-use and anthropogenic datasets to better capture the interactions between weather variability, ecosystem disturbance, and human-driven wildfire processes. The development of a comprehensive wildfire model could improve predictive capabilities and strengthen adaptation strategies in a changing climate. </p></sec><sec id="sec-7"><title>Abbreviations</title><p>TPFTropical Pine Forest</p><p>MRFMontane Rain Forests </p><p>LRFLowland Rain Forests</p><p>FSFFreshwater Swamp Forests</p><p>PSFPeat Swamp Forests</p><p>SSMSunda Shelf Mangroves</p><p>No. Obsnumber of observation</p><p>psd. R2pseudo-R2</p><p>rfrainfall</p><p>tmtemperature</p><p>smsoil moisture</p></sec></body><back><ack><title>Acknowledgements</title><p>The authors gratefully acknowledge the financial support provided by Universitas Gadjah Mada through the Final Project Recognition Grant (<italic>Rekognisi</italic><italic>Tugas</italic><italic> Akhir</italic>) in 2024.</p></ack><sec sec-type="author-contributions"><title>Author Contributions</title><p><bold>Conceptualization</bold>: Asshaffa Naim,  Andung Bayu Sekaranom; <bold>methodology</bold>: Asshaffa Naim; <bold>investigation</bold>:  Asshaffa Naim; <bold>writing—original draft preparation</bold>:  Asshaffa Naim; <bold>writing—review and editing</bold>:  Andung Bayu Sekaranom, Sudrajat,  Fitria Nucifera; <bold>visualization</bold>: Asshaffa Naim. All authors have read and agreed to the published version of the manuscript.</p></sec><sec sec-type="conflict-of-interest"><title>Conflict of Interest</title><p>All authors declare that they have no conflicts of interest.</p></sec><sec sec-type="data-availability"><title>Data Availability</title><p>Data are available upon request.</p></sec><sec sec-type="funding"><title>Funding</title><p>This research was supported by a Grant of Final Project Recognition (Rekognisi Tugas Akhir) by Universitas Gadjah Mada in 2024. The grant was entitled “Analisis Dampak Perubahan Iklim terhadap Kerentanan dan Ketahanan Lingkungan Lokal di Indonesia” with Dr. Sc. 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